Few studies have systematically addressed evolutionary changes in olfactory neuron assemblies, either by genetic drift or as an

adaptation to specific odor environments. We have studied the sense of olfaction in 2 congeneric scarab beetles, Pachnoda

interrupta Olivier and Pachnoda marginata Drury (Coleoptera: Scarabaeidae: Cetoniinae), which are both opportunistic

polyphages, feeding mainly on fruit and flowers. The 2 species occur in dissimilar habitats: P. interrupta is found in dry

savannah, and P. marginata in tropical parts of equatorial Africa. To study how these species may have adapted their sense of

olfaction to their odor environments, we utilized single-unit electrophysiology on olfactory sensilla with a wide selection of

food-related compounds. Despite the differences in habitat, we found that the species shared most of the physiological types

of olfactory receptor neurons (ORNs) encountered, although their proportions frequently varied between the species. The high

degree of conservation in olfaction between the species implies that a similar sensory strategy is efficient for food search in

both habitats. However, shifts in proportions of receptor neuron classes, and slight shifts in response profiles and/or presence

of some ORN classes unique to either species, may reflect adaptation to a different set of hosts.

Key words: comparative study, electrophysiology, olfaction, polyphagous herbivore, single sensillum recordings

Few studies have addressed how olfactory systems may be adapted to different odour environments. I have performed the largest study to date, comparing olfactory receptor neurons in the two congeneric species of scarab beetle, P. marginata and P. interrupta. Both species are fruit- and flower-eaters but geographically separated (equatorial versus sub-Saharan Africa, resp.). They have similar lamellate antennae covered mostly with numerous olfactory sensilla placodea (plate sensilla) and a lesser number of other types, mainly sensilla coeloconica (grooved peg), and smooth peg sensilla of unknown function. By means of single sensillum recordings with tungsten microelectrodes I screened a great number of olfactory sensilla with a large array of odorants. I compared 456 sensilla placodea, containing 212 responding cells, over the whole antennae in both species. The olfactory systems of these two species displayed an amazing degree of conservation, with 20 identified olfactory neuron classes, all except two of which were found in both species with no detectable difference in response profiles. The exceptions were two olfactory receptor neuron classes: methyl benzoate and gamma-nonalactone, which were only found in Pachnoda marginata. In general, the two species showed an almost total overlap in their receptor neuron assemblies. One aim of the study was also to test whether the arrangement of neurons within sensilla was conserved between the species, but this could not be tested as very few combinations of characterized neurons were encountered during the study.

Key words: olfaction, electrophysiology, scarab beetles, Pachnoda marginata, Pachnoda interrupta, single sensillum recording, olfactory receptor neurons

Due to the inherent complexity of decision making in supply chain management (SCM), there is a growing need for modelling supply chain systems with different methodologies. A large number of manufacturing organisations in particular the cement industry in Libya is seeking techniques that can help, identify and implement strategies for developing and improving their supply chain network. Developing a supply chain management system requires the analysis of the manufacturing processes from the initial sourcing to the end customers. The paper reports that the Libyan cement industry has problems related to its productivity. In order to face these problems, cement factories need to increase their production. To accomplish this, they require a clear strategy towards an efficient supply chain.

Auditor independence has received considerable attention in recent

years. This is due to the fact that independently audited financial statements

may result in the generation of true and fair accounting information which will

help stakeholders to form rational expectations about firms and minimise the

agency cost. It can also be argued that lack of independence would lead

auditors to collaborate with the management of firms and would produce

misleading accounting information. Accepting this premise, this study explores

the effects of 12 different variables on the perceptions of auditor independence

in Libya. A sample of five user groups namely; owners, investors, lenders,

managers and auditors were chosen for the survey. The results suggest that all

user groups regard auditor independence as an important factor in forming their

decisions about firms. Amongst other variables, the non-availability of auditing

standards in Libya is found to be the strongest factor which undermines auditor

independence in Libya